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Hippo, Zambezi River, Africa, 1997

The gape-mouthed territorial display of a dominant male hippopotamus, such as this one on the Zambezi River, looks more like a colossal yawn. During mating season, males will claim a length of shoreline and fiercely defend it, occasionally even killing a bachelor rival.

(Photo shot on assignment for, but not published in, "Down the Zambezi," October 1997, National Geographic magazine)

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Photograph by Chris Johns